They found that the genetic sequences of the Tuscan men varied significantly from those of men in surrounding regions in Italy, and that the men from Murlo and Volterra were the most closely related to men from Turkey.
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He finds Etruscan on one hand genetically related to the Rhaetic language spoken in the Alps north of Etruria, suggesting autochthonous connections, but on the other hand the Lemnian language found on the "Lemnos stele" is closely related to Etruscan, entailing either Etruscan presence in "Tyrsenian" Lemnos, or "Tyrsenian" expansion westward to Etruria.
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