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He was the son of Claude Louis Hector de Villars and of Jeanne Angélique Rocque de Varengeville, and the grandson of Pierre de Villars.
D'Arco was born in Arco, Trentino, and already had 30 years distinguished service when he was ordered by Louis William, Margrave of Baden-Baden in 1703 to defend Breisach to the last man against a French attack under Villars.
Six marshals of France have been given the even more exalted rank of "marshal general of France" (maréchal général de France): Biron, Lesdiguières, Turenne, Villars, Saxe and Soult.